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I think its both.
The koi "work the bottom" constantly seiving mud to get the "good stuff" out of it. I beleive they iingest lots of minerals like this. They also like to stick thier snouts into crawfish holes and pump the crawfish out with thier gills. Watching this behavior is quite interesting.
I beleive the koi smells a crawfish that has recently molted, they like softshells just like I do, but I want mine fried.Sometimes several koi will work one area, each with its head in the side of the pond. A big plume of really muddy water will be all around them. Unfortunately this behavior is really hard on the pond banks, undermining them and causing the tractor to fall into the pond on many occasions. The ponds must be reshaped every few years (as is done in Japan).
My koi get an amazing diversity of live foods, these change with the seasons and they feed heavily upon whatever is in abundance at the time. One favorite is mayflies. On the mornig after a big hatch, the yellow mayflies will be all over he pond's surface with the koi slurping them up, gorging until they are all gone.
On a windy summer's day they'll be right at the upwind edge waiting for a tasty morsel to blow in. A grasshopper is a favorite, but grass seeds, and other stuff is also eaten as it blows in.
A blustery early spring day finds the koi at the other end of the pond as the wind is now from the North and blowing in caterpillars from the trees. They eat the caterpillars as they hit the water.
A big rainstorm in late summer washes earthworms into the ponds by the thousands, koi there waiting as they wash in.
In most of my ponds, only about half of what the fish get to eat comes out of the feed bag.
Both the mud pond environment with its minerals and water quality as well as its natural foods contributes to the development of the koi.
However, koi do not finish up good in the mud. They need the closed environemtn in order to be brought fully into show form. Hence the need for both environments.
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