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Now, I figure that Aquitori is asking about final size of the offspring and not about "fry size" as it would be quite obvious that the male would not in any way, shape, form, or fashion have ANY influence on the size of his spawn's fry. And Aquitori does talk of the parents (plural).
And Dick answered that question...but one other dynamic needs to be mentioned...there are many strategies within a species dna that dictates the method used to enhance the odds that the offspring ( a number of the offspring suitable to maintain the continuance of the individual's involved DNA make it through to yet another generation) survive to adulthood.
The strategy utilized by carp is to produce an astonding number of offspring. One of the dynamics which increases the likelihood that the needed number of offspring (just a few) will reach adulthood is to have an extreme variation in the genetic make-up of the offspring......Sorta like quick picks for the lottery. And in carp the variation doesn't look that great to us and it isn't because all the odd balls and their genes are weeded out by predators and such...
But then man steps in and saves the really odd-looking fish. And when those are bred the genes available are extreme and the "mixing" still occurs...and the whole gambit is thrown....and that explains why the "culling ain't gonna be able to be stopped. Koi do not throw true to what they look like to any great extent.
The same thing happens when big parents are bred...the genes to throw "big koi" are passed on..just not equally to all the spawn...chances are it will be skewed towards the behavior, metabolism,style, color, shape, and length of the parents....but remember the way this animal has been able to survive all sorts of enviroments for millions of years from one generation to the next is by making all the genetic options possible.
The nurture(physical) aspect entertwines with the nature (genes) aspect and it lets the ones with the right genes for that environment survive, grow, mature, and breed.
Yes simplified...
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